הזיהויים של onotole

These appeared in the microscopy of the lichen I was looking at — my previous observation, Platismata tuckermanii — but I have not ever seen something like this before and did not see it mentioned in the Brodo lichen book. It really did look “a part of” the lichen but maybe it is something deposited in/on the lichen, eggs? Not sure. Central sphere seems to have many smaller spheres in them. They are large.

What are they?